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At this writing, public databases contain the completed, contiguous sequences of a large number of bacterial genomes (e.g.,) (
1
), the yeast
Saccharomyces cerevisiae
genome (
2
); and the genomes of the nematode (
3
), Drosophila (
4
), and Arabidopsis (
5
). Public sequencing projects for many other genomes, including several large ones, are in progress. In the case of the publicly funded human genome sequencing effort, more than 45% of the >3-Gb genome is in finished form, including several completed chromosomes, with the remaining 55% expected to be finished by the spring of 2003. The term
finished
to describe sequence has special meaning and significance in the public genome-sequencing arena, where there is a general agreement that a large-insert clone, chromosome, or genome is considered finished when the accuracy of the sequence exceeds 99.99% (i.e., <1 error in 10,000 bp) and all gaps that can be filled by known techniques are filled, so that the sequence is either completely contiguous or has very few, well-annotated gaps.
